It's a bit complicated topology..
e.g. U3-2 is wired as non-inverting configuration with
its own stage gain of (1 + R19/R21) BUT actuall signal comming into pin5 is what comes out of U3-1 divided with R22/[(R20//(R18+R9+R8)] (or multiplied with the reverse fraction - to be more true-blue voltage divider formula..)
U1-2, U2-1, U2-2 & U3-1 are wired as inverting configuration with unity gain BUT their non-inverting inputs are biased via LDRs and toghether with respective capacitors (C5, C7, C8 & C9) high-pass filter configuration form
all-pass filters..
At high frequencies caps are shorts so we have unity gain voltage buffers (no phase lead)..
At low frequencies caps are open circuits and we have unity gain inverting amplifiers (180
o phase lead)
At corner frequency of HPFs (0.159/R*C) we have 90
o phase lead..
(I let you guess the reason for it..

hint: at corner frequency LDRs resistance and capacitors impedance are equal..)
LDRs resistance varies with light intensity which variation is militated by lower right circuitry (LFO) resulting into LFO frequency depended "moving" phase lead..