You must have a supply issue around IC2..
Check R30/R31 values/ratio or better check VR,,
(it should measure about 60% of VC..)
Q1 to Q3 are bottomed due to high Source voltage caused by high Gate voltage..
(they should be biased at S: 1/3VC and D: 2/3VC..)
As for Q4, you should be able to bias it (via 100k trimpot) at S: 1/4VC and D: 3/4VC..